In a seemingly utopian world powered by Mana, a miraculous technological energy, humanity lives free of war and poverty. However, a small percentage of people born without Mana are labeled "Norma," treated as subhuman outcasts, and exiled. Angeleze Ikaruga Misurugi, the beloved crown princess of the Misurugi Empire, has her life shattered during her sixteenth-birthday coronation when her brother, Algrus, reveals she is actually a Norma. Stripped of her title and royal status, Ange is banished to Arzenal, a military base where Norma are forced to pilot mechs called Paramails to fight monstrous interdimensional invaders known as Dragons.Upon arrival, the defiant and arrogant Ange struggles to adapt, refusing to accept her new reality or fit in with the hardened, cynical Norma soldiers, particularly Salia and Hilda. During her first brutal battle, Ange experiences profound trauma as her squadmates are slaughtered, destroying her naive worldview. Touched by the harshness of Arzenal and the death of her commanding officer, Jill—a rebel leader plotting against the ruling order—Ange undergoes a drastic personality shift, hardening into a ruthless, pragmatic survivor who learns to pilot the custom Paramail, Vilkiss.Plot twists escalate when Ange discovers that Mana is a parasitic energy created by Embryo, a godlike being who controls humanity through a continuous cycle of reincarnation. The world leaders, including Ange’s former fiancé Julius, are complicit in this grand deception. During a chaotic battle, Ange and a mysterious
Dragon warrior named Tusk are thrown through dimensional rifts, stranding them in an alternate, desolate Earth. There, Ange learns the truth about the Dragons: they are actually mutated humans who rebelled against Embryo's tyranny centuries ago.Returning to their home dimension, Ange, Jill, and the awakened rebel Norma wage a desperate war against Embryo, who attempts to enact "Rondo," a plan to reset the universe and make Ange his ultimate bride. Along the way, Ange mends fences with her former enemies, uniting the Norma, the Dragons, and disillusioned Mana-users. The dramatic climax culminates in a fierce final battle against Embryo. Ange sacrifices her former royal identity entirely, utilizing the full, latent power of the Vilkiss. Ultimately, Ange defeats Embryo, destroying the false paradise of Mana and liberating humanity from its cycle of control. In
the aftermath, Ange and Tusk choose a life of freedom, riding off into a hopeful new dawn together with their surviving comrades, ready to build a genuinely free world from the ashes of the old one.