The narrative kicks off with the charismatic and hot-headed Yuuto Sakurai, the temporal warrior known as Kamen Rider Zeronos, finding himself in the middle of a bizarre anomaly that threatens the very fabric of the DenLiner and the timelines. Accompanied by his loyal, perpetually hungry Imagin partner, Deneb, Yuuto is driven by a fierce motivation to protect the flow of time and, more personally, the memories that anchor his existence in the present. However, a strange temporal distortion begins to sap Yuuto’s remaining memories, directly weakening his physical manifestation and his ability to transform into Zeronos. Complicating matters further, a rogue group of New Imagin appears, seeking to exploit this temporal vulnerability to rewrite history for their own chaotic ends. Amidst this high-stakes chaos, Airi Sakurai—the woman Yuuto deeply cares for and is destined to protect—becomes unwittingly entangled in the plot, serving as the emotional anchor for Yuuto's relentless drive. Ryotaro Nogami, the original Kamen Rider Den-O, steps in to offer assistance alongside his own diverse roster of Imagin, but the sheer volatility of Yuuto’s
fading memories creates a dangerous rift between the heroes. Yuuto must confront not only the physical threat of the New Imagin but also the agonizing prospect of being completely forgotten by the world he fights to save. The emotional crux of the story hinges on Yuuto's internal struggle, balancing his duty as a protector with his fading humanity. Deneb goes to great lengths, distributing his signature "Deneb Candy" and trying to mend fences, but the crisis escalates as the primary antagonist reveals a master plan to sever Yuuto's tie to Airi permanently. This revelation sparks a fierce determination in Yuuto, overriding his fading condition with sheer willpower. The climax erupts into an explosive showdown aboard a derailed timeline, where Yuuto summons every ounce of his remaining memory to power the Zeronos Belt. Transforming into his fiery Vega Form, he delivers a decisive, spectacular defeat to the rogue Imagin. In
the aftermath, the timeline stabilizes, and while Yuuto's memories remain precariously thin, a poignant moment with Airi reaffirms his place in the timeline.